"Mushi": the most basic forms of life in the world. They exist without any goals or purposes aside from simply "being." They are beyond the shackles of the words "good" and "evil." Mushi can exist in countless forms and are capable of mimicking things from the natural world such as plants, diseases, and even phenomena like rainbows. This is, however, just a vague definition of these entities that inhabit the vibrant world of Mushishi, as to even call them a form of life would be an oversimplification. Detailed information on Mushi is scarce because the majority of humans are unaware of their existence. So what are Mushi and why do they exist? This is the question that a "Mushishi," Ginko, ponders constantly. Mushishi are those who research Mushi in hopes of understanding their place in the world's hierarchy of life. Ginko chases rumors of occurrences that could be tied to Mushi, all for the sake of finding an answer. It could, after all, lead to the meaning of life itself. [Written by MAL Rewrite]

Now in his second year of high school, Rei Kiriyama continues pushing through his struggles in the professional shogi world as well as his personal life. Surrounded by vibrant personalities at the shogi hall, the school club, and in the local community, his solitary shell slowly begins to crack. Among them are the three Kawamoto sisters—Akari, Hinata, and Momo—who forge an affectionate and familial bond with Rei. Through these ties, he realizes that everyone is burdened by their own emotional hardships and begins learning how to rely on others while supporting them in return. Nonetheless, the life of a professional is not easy. Between tournaments, championships, and title matches, the pressure mounts as Rei advances through the ranks and encounters incredibly skilled opponents. As he manages his relationships with those who have grown close to him, the shogi player continues to search for the reason he plays the game that defines his career. [Written by MAL Rewrite]

Chihaya Ayase, a strong-willed and tomboyish girl, grows up under the shadow of her older sister. With no dreams of her own, she is contented with her share in life till she meets Arata Wataya. The quiet transfer student in her elementary class introduces her to competitive karuta, a physically and mentally demanding card game inspired by the classic Japanese anthology of Hundred Poets. Captivated by Arata's passion for the game and inspired by the possibility of becoming the best in Japan, Chihaya quickly falls in love with the world of karuta. Along with the prodigy Arata and her haughty but hard-working friend Taichi Mashima, she joins the local Shiranami Society. The trio spends their idyllic childhood days playing together, until circumstances split them up. Now in high school, Chihaya has grown into a karuta freak. She aims to establish the Municipal Mizusawa High Competitive Karuta Club, setting her sights on the national championship at Omi Jingu. Reunited with the now indifferent Taichi, Chihaya's dream of establishing a karuta team is only one step away from becoming true: she must bring together members with a passion for the game that matches her own. [Written by MAL Rewrite]

Fourteen-year-old Yukio "Koyuki" Tanaka is a dispirited young boy with no goals in life. However, this all changes when Koyuki saves a strange-looking dog named Beck from being harassed by a group of local kids. The dog's owner, 16-year-old Ryuusuke "Ray" Minami, is an emerging guitarist and the former member of a popular rock band. After Koyuki meets Ray again in a diner, the older boy leads him to his former band's meeting place and dazzles Koyuki with his amazing guitar skills. Slowly becoming interested in the glamour of western rock culture, Koyuki decides to start playing the guitar while helping Ray achieve his dream of leading the ultimate rock band. Together with Ray's younger sister Maho and a few other members, the two boys launch their career into the world of rock by forming a band called BECK. Beck follows the group's struggles and successes as they spread their fame across Japan. [Written by MAL Rewrite]